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Naran, Daksha. Rhodes University; 1997. Cytogenetic studies of Pseudobarbus and selected Barbus (Pisces: Cyprinidae) of southern Africa.
Naran, Daksha, Skelton, Paul H, Villet, Martin H. Karyology of three evolutionarily hexaploid southern African species of yellowfish, Labeobarbus Rüppel, 1836 (Cyprinidae). 2007.
Naran, Daksha, Skelton, Paul H, Villet, Martin H. African Zoology; 2006. Karyology of the redfin minnows, genus Pseudobarbus Smith, 1841 (Teleostei: Cyprinidae): one of the evolutionarily tetraploid lineages of South African barbines.